to be prudent; to be on one's guard, alert; to be clever, wise, of good understanding James Lockhart, Nahuatl as Written: Lessons in Older Written Nahuatl, with Copious Examples and Texts (Stanford: Stanford University Press and UCLA Latin American Studies, 2001), 220.
mimatcatlaloa = to they run prudently (combines imati and tlaloa with the connector "ca," which refers to the manner in which the second verb is done) Rebecca Horn's notes from Nahuatl classes with James Lockhart at UCLA.